At what scale does environmental variation promote species diversity?
How does variation between individuals influence extinction risk and the maintenance of species diversity?
Freshwater ecosystems are the most threatened on earth and the impacts continue to increase. How will they respond?

I combine theory and experiments to explore the generation, maintenance, and loss of diversity.

The first major component of my research explores the influence of scale-dependent environmental heterogeneity on species coexistence.

The second major component of my research explores the influence of dynamics occurring at the level of individuals – or individual variation – on species coexistence and extinction risk.

I address general questions and have therefore worked in a wide variety of systems, including subtidal marine environments, terrestrial plant communities, and freshwater ecosystems. For the last five years my research has focused exclusively on the ecology and evolutionary biology of plants.

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