At what scale does environmental variation promote species diversity?
How does variation between individuals influence extinction risk and the maintenance of species diversity?
Freshwater ecosystems are the most threatened on earth and the impacts continue to increase. How will they respond?

I combine theory and experiments to explore the generation, maintenance, and loss of species (and genetic) diversity.

The first major component of my research explores the ecological and evolutionary effects of differences between individuals on species coexistence and extinction risk.

The second major component of my research explores the influence of scale-dependent environmental heterogeneity on species coexistence.

I address general questions and have therefore worked in a wide variety of systems, including animal and plant communities in terrestrial, freshwater, and marine environments.

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